/* ============================================================
   MERAMEC BIO EVAL CENTER — Design System
   Based on Zapier-inspired design from DESIGN.md
   ============================================================ */

/* --- CSS Custom Properties --- */
:root {
  /* Colors */
  --color-bg: #fffefb;
  --color-bg-alt: #fffdf9;
  --color-text: #201515;
  --color-text-body: #36342e;
  --color-text-muted: #939084;
  --color-accent: #ff4f00;
  --color-border: #c5c0b1;
  --color-border-strong: #36342e;
  --color-surface: #eceae3;
  --color-surface-mid: #b5b2aa;
  --color-dark: #201515;
  --color-light: #fffefb;

  /* Fonts */
  --font-display: 'Plus Jakarta Sans', 'Degular Display', system-ui, sans-serif;
  --font-primary: 'Inter', Helvetica, Arial, sans-serif;
  --font-editorial: 'Cormorant', 'GT Alpina', Georgia, serif;

  /* Spacing */
  --space-1: 4px;
  --space-2: 8px;
  --space-3: 12px;
  --space-4: 16px;
  --space-5: 20px;
  --space-6: 24px;
  --space-8: 32px;
  --space-10: 40px;
  --space-12: 48px;
  --space-14: 56px;
  --space-16: 64px;
  --space-20: 80px;

  /* Radius */
  --radius-tight: 3px;
  --radius-sm: 4px;
  --radius-md: 5px;
  --radius-lg: 8px;
  --radius-pill: 20px;

  /* Container */
  --container-max: 1200px;
  --container-padding: 24px;
}

/* --- Reset & Base --- */
*, *::before, *::after {
  box-sizing: border-box;
  margin: 0;
  padding: 0;
}

html {
  font-size: 16px;
  -webkit-font-smoothing: antialiased;
  -moz-osx-font-smoothing: grayscale;
  scroll-behavior: smooth;
}

body {
  font-family: var(--font-primary);
  font-weight: 400;
  line-height: 1.25;
  color: var(--color-text-body);
  background-color: var(--color-bg);
  letter-spacing: -0.16px;
}

img {
  max-width: 100%;
  height: auto;
  display: block;
  background-color: var(--color-surface);
}

a {
  color: var(--color-text);
  text-decoration: underline;
  transition: all 0.2s ease;
}

a:hover {
  text-decoration: none;
}

ul, ol {
  list-style: none;
}

h1, h2, h3, h4, h5, h6 {
  color: var(--color-text);
  font-family: var(--font-primary);
  font-weight: 500;
}

/* --- Typography --- */
.display-hero-xl {
  font-family: var(--font-display);
  font-size: 5rem;
  font-weight: 500;
  line-height: 0.90;
  color: var(--color-text);
}

.display-hero {
  font-family: var(--font-display);
  font-size: 3.5rem;
  font-weight: 500;
  line-height: 0.90;
  color: var(--color-text);
}

.display-hero-sm {
  font-family: var(--font-display);
  font-size: 2.5rem;
  font-weight: 500;
  line-height: 0.90;
  color: var(--color-text);
}

.section-heading {
  font-family: var(--font-primary);
  font-size: 3rem;
  font-weight: 500;
  line-height: 1.04;
  color: var(--color-text);
}

.editorial-heading {
  font-family: var(--font-editorial);
  font-size: 3rem;
  font-weight: 250;
  letter-spacing: -1.92px;
  color: var(--color-text);
}

.editorial-sub {
  font-family: var(--font-editorial);
  font-size: 2.5rem;
  font-weight: 300;
  line-height: 1.08;
  letter-spacing: -1.6px;
  color: var(--color-text);
}

.sub-heading-lg {
  font-size: 2.25rem;
  font-weight: 500;
  letter-spacing: -1px;
}

.sub-heading {
  font-size: 2rem;
  font-weight: 400;
  line-height: 1.25;
}

.sub-heading-md {
  font-size: 1.75rem;
  font-weight: 500;
}

.card-title {
  font-size: 1.5rem;
  font-weight: 600;
  letter-spacing: -0.48px;
  color: var(--color-text);
}

.body-large {
  font-size: 1.25rem;
  font-weight: 400;
  line-height: 1.20;
  letter-spacing: -0.2px;
  color: var(--color-text-body);
}

.body-emphasis {
  font-size: 1.125rem;
  font-weight: 600;
  line-height: 1.00;
}

.label-upper {
  font-size: 0.875rem;
  font-weight: 600;
  text-transform: uppercase;
  letter-spacing: 0.5px;
  color: var(--color-text-muted);
}

.micro {
  font-size: 0.75rem;
  font-weight: 600;
  letter-spacing: 0.5px;
  text-transform: uppercase;
}

/* --- Layout --- */
.container {
  max-width: var(--container-max);
  margin: 0 auto;
  padding-left: var(--container-padding);
  padding-right: var(--container-padding);
}

.section {
  padding: var(--space-16) 0;
}

.section--sm {
  padding: var(--space-10) 0;
}

.section--lg {
  padding: var(--space-20) 0;
}

.section--cream {
  background-color: var(--color-bg);
}

.section--alt {
  background-color: var(--color-bg-alt);
}

.section--dark {
  background-color: var(--color-dark);
  color: var(--color-light);
}

.section--dark h2,
.section--dark h3,
.section--dark .card-title,
.section--dark .section-heading {
  color: var(--color-light);
}

.section--dark .body-large,
.section--dark p {
  color: var(--color-border);
}

.section__border-top {
  border-top: 1px solid var(--color-border);
}

.grid {
  display: grid;
  gap: var(--space-6);
}

.grid--2 {
  grid-template-columns: repeat(2, 1fr);
}

.grid--3 {
  grid-template-columns: repeat(3, 1fr);
}

.grid--4 {
  grid-template-columns: repeat(4, 1fr);
}

.flex {
  display: flex;
}

.flex--center {
  align-items: center;
}

.flex--between {
  justify-content: space-between;
}

.flex--wrap {
  flex-wrap: wrap;
}

.flex--gap-2 {
  gap: var(--space-2);
}

.flex--gap-3 {
  gap: var(--space-3);
}

.flex--gap-4 {
  gap: var(--space-4);
}

.flex--gap-6 {
  gap: var(--space-6);
}

.text-center {
  text-align: center;
}

.text-muted {
  color: var(--color-text-muted);
}

.mt-2 { margin-top: var(--space-2); }
.mt-3 { margin-top: var(--space-3); }
.mt-4 { margin-top: var(--space-4); }
.mt-6 { margin-top: var(--space-6); }
.mt-8 { margin-top: var(--space-8); }
.mt-10 { margin-top: var(--space-10); }
.mt-12 { margin-top: var(--space-12); }
.mb-2 { margin-bottom: var(--space-2); }
.mb-3 { margin-bottom: var(--space-3); }
.mb-4 { margin-bottom: var(--space-4); }
.mb-6 { margin-bottom: var(--space-6); }
.mb-8 { margin-bottom: var(--space-8); }
.mb-10 { margin-bottom: var(--space-10); }
.max-w-prose { max-width: 720px; margin-left: auto; margin-right: auto; }

/* --- Navigation --- */
.nav {
  position: sticky;
  top: 0;
  z-index: 100;
  background-color: var(--color-bg);
  border-bottom: 1px solid var(--color-border);
}

.nav__inner {
  display: flex;
  align-items: center;
  justify-content: space-between;
  max-width: var(--container-max);
  margin: 0 auto;
  padding: var(--space-4) var(--container-padding);
}

.nav__logo {
  font-family: var(--font-display);
  font-size: 1.25rem;
  font-weight: 700;
  color: var(--color-text);
  text-decoration: none;
  display: flex;
  align-items: center;
  gap: var(--space-2);
  white-space: nowrap;
}

.nav__logo:hover {
  text-decoration: none;
}

.nav__logo-mark {
  display: inline-flex;
  align-items: center;
  justify-content: center;
  width: 36px;
  height: 36px;
  background-color: var(--color-accent);
  color: var(--color-light);
  border-radius: var(--radius-sm);
  font-weight: 700;
  font-size: 1.125rem;
  font-family: var(--font-display);
}

.nav__links {
  display: flex;
  align-items: center;
  gap: var(--space-1);
}

.nav__link {
  font-size: 0.9375rem;
  font-weight: 500;
  color: var(--color-text);
  text-decoration: none;
  padding: var(--space-2) var(--space-3);
  border-radius: var(--radius-sm);
  transition: background-color 0.15s ease;
}

.nav__link:hover {
  background-color: var(--color-surface);
  text-decoration: none;
}

.nav__link--active {
  box-shadow: rgb(255, 79, 0) 0px -4px 0px 0px inset;
}

.nav__cta {
  margin-left: var(--space-4);
}

.nav__mobile-toggle {
  display: none;
  background: none;
  border: 1px solid var(--color-border);
  border-radius: var(--radius-sm);
  padding: var(--space-2) var(--space-3);
  font-size: 1.25rem;
  cursor: pointer;
  color: var(--color-text);
  line-height: 1;
}

.nav__mobile-menu {
  display: none;
  position: absolute;
  top: 100%;
  left: 0;
  right: 0;
  background-color: var(--color-bg);
  border-bottom: 1px solid var(--color-border);
  padding: var(--space-4) var(--container-padding) var(--space-6);
}

.nav__mobile-menu--open {
  display: block;
}

.nav__mobile-menu .nav__link {
  display: block;
  padding: var(--space-3) 0;
  border-bottom: 1px solid var(--color-surface);
}

.nav__mobile-menu .btn {
  margin-top: var(--space-4);
  width: 100%;
  text-align: center;
}

/* --- Hero --- */
.hero {
  padding: var(--space-16) 0 var(--space-12);
  border-bottom: 1px solid var(--color-border);
}

.hero--home {
  padding: var(--space-20) 0 var(--space-16);
}

.hero--home .hero__title {
  font-size: 5rem;
}

.hero__content {
  max-width: 900px;
}

.hero__content--center {
  margin: 0 auto;
  text-align: center;
}

.hero__content--center .hero__subtitle {
  margin-left: auto;
  margin-right: auto;
}

.hero__content--center .hero__actions {
  justify-content: center;
}

.hero__content--center .breadcrumb {
  justify-content: center;
}

/* Home hero with background image */
.hero--home {
  background-image: linear-gradient(rgba(255, 254, 251, 0.88), rgba(255, 254, 251, 0.92)), url('https://images.unsplash.com/photo-1532094349884-543bc11b234d?w=1600&h=900&fit=crop&auto=format&q=80');
  background-size: cover;
  background-position: center;
}

.hero__label {
  font-size: 0.875rem;
  font-weight: 600;
  text-transform: uppercase;
  letter-spacing: 0.5px;
  color: var(--color-accent);
  margin-bottom: var(--space-4);
}

.hero__title {
  font-family: var(--font-display);
  font-size: 3.5rem;
  font-weight: 500;
  line-height: 0.90;
  color: var(--color-text);
  margin-bottom: var(--space-6);
}

.hero__subtitle {
  font-size: 1.25rem;
  font-weight: 400;
  line-height: 1.40;
  color: var(--color-text-body);
  max-width: 640px;
}

.hero__actions {
  display: flex;
  gap: var(--space-4);
  margin-top: var(--space-8);
  flex-wrap: wrap;
}

/* --- Breadcrumb --- */
.breadcrumb {
  display: flex;
  align-items: center;
  gap: var(--space-2);
  margin-bottom: var(--space-6);
  font-size: 0.875rem;
  color: var(--color-text-muted);
}

.breadcrumb__link {
  color: var(--color-text-muted);
  text-decoration: none;
}

.breadcrumb__link:hover {
  color: var(--color-accent);
  text-decoration: none;
}

.breadcrumb__separator {
  color: var(--color-border);
}

.breadcrumb__current {
  color: var(--color-text);
  font-weight: 500;
}

/* --- Buttons --- */
.btn {
  display: inline-flex;
  align-items: center;
  justify-content: center;
  gap: var(--space-2);
  font-family: var(--font-primary);
  font-weight: 600;
  font-size: 1rem;
  text-decoration: none;
  cursor: pointer;
  transition: all 0.2s ease;
  border: 1px solid transparent;
  line-height: 1;
}

.btn:hover {
  text-decoration: none;
}

.btn--primary {
  background-color: var(--color-accent);
  color: var(--color-light);
  padding: 8px 16px;
  border-radius: var(--radius-sm);
  border-color: var(--color-accent);
}

.btn--primary:hover {
  background-color: #e64600;
  border-color: #e64600;
  color: var(--color-light);
}

.btn--dark {
  background-color: var(--color-dark);
  color: var(--color-light);
  padding: 20px 24px;
  border-radius: var(--radius-lg);
  border-color: var(--color-dark);
}

.btn--dark:hover {
  background-color: var(--color-border);
  color: var(--color-text);
  border-color: var(--color-border);
}

.btn--light {
  background-color: var(--color-surface);
  color: var(--color-text-body);
  padding: 20px 24px;
  border-radius: var(--radius-lg);
  border-color: var(--color-border);
}

.btn--light:hover {
  background-color: var(--color-border);
  color: var(--color-text);
}

.btn--pill {
  background-color: var(--color-bg);
  color: var(--color-text-body);
  padding: 0 16px;
  height: 36px;
  border-radius: var(--radius-pill);
  border-color: var(--color-border);
}

.btn--pill:hover {
  background-color: var(--color-surface);
}

.btn--sm {
  font-size: 0.875rem;
  padding: 8px 16px;
}

.btn--lg {
  font-size: 1rem;
  padding: 20px 32px;
}

.btn--arrow::after {
  content: " →";
  transition: transform 0.2s ease;
}

.btn--arrow:hover::after {
  transform: translateX(2px);
}

/* --- Cards --- */
.card {
  background-color: var(--color-bg);
  border: 1px solid var(--color-border);
  border-radius: var(--radius-md);
  overflow: hidden;
  transition: border-color 0.2s ease;
}

.card:hover {
  border-color: var(--color-surface-mid);
}

.card__image {
  aspect-ratio: 16 / 10;
  overflow: hidden;
  border-bottom: 1px solid var(--color-border);
  background-color: var(--color-surface);
}

.card__image img {
  width: 100%;
  height: 100%;
  object-fit: cover;
}

.card__image--placeholder {
  display: flex;
  align-items: center;
  justify-content: center;
  font-size: 2.5rem;
  color: var(--color-text-muted);
  background: linear-gradient(135deg, var(--color-surface) 0%, var(--color-bg-alt) 100%);
}

.card__body {
  padding: var(--space-6);
}

.card__label {
  font-size: 0.75rem;
  font-weight: 600;
  text-transform: uppercase;
  letter-spacing: 0.5px;
  color: var(--color-accent);
  margin-bottom: var(--space-2);
}

.card__title {
  font-size: 1.5rem;
  font-weight: 600;
  letter-spacing: -0.48px;
  color: var(--color-text);
  margin-bottom: var(--space-3);
}

.card__title a {
  color: inherit;
  text-decoration: none;
}

.card__title a:hover {
  color: var(--color-accent);
}

.card__text {
  font-size: 1rem;
  line-height: 1.50;
  color: var(--color-text-body);
  margin-bottom: var(--space-4);
}

.card__meta {
  font-size: 0.8125rem;
  color: var(--color-text-muted);
  display: flex;
  gap: var(--space-3);
  flex-wrap: wrap;
}

.card__meta span {
  display: inline-flex;
  align-items: center;
  gap: 4px;
}

.card__cta {
  display: inline-flex;
  align-items: center;
  gap: 4px;
  font-size: 0.9375rem;
  font-weight: 600;
  color: var(--color-accent);
  text-decoration: none;
  margin-top: var(--space-4);
}

.card__cta:hover {
  text-decoration: none;
  gap: 8px;
}

.card__cta::after {
  content: "→";
  transition: transform 0.2s ease;
}

.card--horizontal {
  display: grid;
  grid-template-columns: 280px 1fr;
}

.card--horizontal .card__image {
  aspect-ratio: auto;
  border-bottom: none;
  border-right: 1px solid var(--color-border);
}

.card--featured {
  border-radius: var(--radius-lg);
}

/* --- Stats --- */
.stats {
  display: flex;
  gap: var(--space-10);
  flex-wrap: wrap;
}

.stats--centered {
  justify-content: center;
}

.stats__item {
  text-align: center;
  display: flex;
  flex-direction: column;
  align-items: center;
}

.stats__number {
  display: block;
  font-family: var(--font-primary);
  font-size: 3rem;
  font-weight: 500;
  color: var(--color-text);
  line-height: 1;
}

.stats__label {
  display: block;
  font-size: 0.9375rem;
  color: var(--color-text-muted);
  margin-top: var(--space-2);
}

/* --- Trust Bar --- */
.trust-bar {
  display: flex;
  align-items: center;
  justify-content: center;
  gap: var(--space-10);
  flex-wrap: wrap;
  padding: var(--space-8) 0;
}

.trust-bar__item {
  display: flex;
  align-items: center;
  gap: var(--space-2);
  font-size: 0.875rem;
  font-weight: 600;
  color: var(--color-text-muted);
  text-transform: uppercase;
  letter-spacing: 0.5px;
}

.trust-bar__badge {
  display: inline-flex;
  align-items: center;
  justify-content: center;
  width: 48px;
  height: 48px;
  border: 2px solid var(--color-accent);
  border-radius: var(--radius-lg);
  font-size: 0.625rem;
  font-weight: 700;
  color: var(--color-accent);
  background-color: var(--color-bg);
  letter-spacing: 0.3px;
}

/* --- Filter --- */
.filter-bar {
  display: flex;
  gap: var(--space-2);
  flex-wrap: wrap;
  margin-bottom: var(--space-8);
}

.filter-btn {
  display: inline-flex;
  align-items: center;
  padding: 0 16px;
  height: 36px;
  font-family: var(--font-primary);
  font-size: 0.875rem;
  font-weight: 500;
  color: var(--color-text-body);
  background-color: var(--color-bg);
  border: 1px solid var(--color-border);
  border-radius: var(--radius-pill);
  cursor: pointer;
  transition: all 0.15s ease;
}

.filter-btn:hover {
  background-color: var(--color-surface);
}

.filter-btn--active {
  background-color: var(--color-text);
  color: var(--color-light);
  border-color: var(--color-text);
}

.filter-btn--active:hover {
  background-color: var(--color-text-body);
}

/* --- Forms --- */
.form__group {
  margin-bottom: var(--space-5);
}

.form__label {
  display: block;
  font-size: 0.875rem;
  font-weight: 600;
  color: var(--color-text);
  margin-bottom: var(--space-2);
}

.form__input,
.form__select,
.form__textarea {
  width: 100%;
  font-family: var(--font-primary);
  font-size: 1rem;
  color: var(--color-text);
  background-color: var(--color-bg);
  border: 1px solid var(--color-border);
  border-radius: var(--radius-md);
  padding: var(--space-3) var(--space-4);
  transition: border-color 0.2s ease;
}

.form__input:focus,
.form__select:focus,
.form__textarea:focus {
  outline: none;
  border-color: var(--color-accent);
}

.form__input::placeholder,
.form__textarea::placeholder {
  color: var(--color-text-muted);
}

.form__textarea {
  min-height: 120px;
  resize: vertical;
}

.form__select {
  appearance: none;
  background-image: url("data:image/svg+xml,%3Csvg xmlns='http://www.w3.org/2000/svg' width='12' height='8' fill='%23939084'%3E%3Cpath d='M6 8L0 0h12z'/%3E%3C/svg%3E");
  background-repeat: no-repeat;
  background-position: right 16px center;
  padding-right: 40px;
}

.form__row {
  display: grid;
  grid-template-columns: 1fr 1fr;
  gap: var(--space-5);
}

/* --- Timeline --- */
.timeline {
  position: relative;
  padding-left: var(--space-12);
}

.timeline::before {
  content: "";
  position: absolute;
  left: 16px;
  top: 0;
  bottom: 0;
  width: 2px;
  background-color: var(--color-border);
}

.timeline__item {
  position: relative;
  padding-bottom: var(--space-8);
}

.timeline__item:last-child {
  padding-bottom: 0;
}

.timeline__marker {
  position: absolute;
  left: calc(-1 * var(--space-12) + 3px);
  top: 2px;
  width: 28px;
  height: 28px;
  border-radius: 50%;
  background-color: var(--color-accent);
  border: 3px solid var(--color-bg);
  box-shadow: 0 0 0 2px var(--color-border);
  display: flex;
  align-items: center;
  justify-content: center;
}

.timeline__step {
  font-size: 0.6875rem;
  font-weight: 700;
  color: var(--color-light);
  letter-spacing: 0;
  margin-bottom: 0;
}

.timeline__title {
  font-size: 1.25rem;
  font-weight: 600;
  color: var(--color-text);
  margin-bottom: var(--space-2);
}

.timeline__text {
  font-size: 1rem;
  line-height: 1.50;
  color: var(--color-text-body);
}

/* --- Blog / Sidebar Layout --- */
.blog-layout {
  display: grid;
  grid-template-columns: 1fr 320px;
  gap: var(--space-10);
  align-items: start;
}

.blog-sidebar {
  position: sticky;
  top: 80px;
}

.blog-sidebar__section {
  margin-bottom: var(--space-8);
}

.blog-sidebar__title {
  font-size: 0.875rem;
  font-weight: 600;
  text-transform: uppercase;
  letter-spacing: 0.5px;
  color: var(--color-text);
  margin-bottom: var(--space-4);
  padding-bottom: var(--space-3);
  border-bottom: 1px solid var(--color-border);
}

.blog-sidebar__link {
  display: block;
  padding: var(--space-2) 0;
  font-size: 0.9375rem;
  color: var(--color-text-body);
  text-decoration: none;
  transition: color 0.15s ease;
}

.blog-sidebar__link:hover {
  color: var(--color-accent);
  text-decoration: none;
}

.blog-sidebar__link--active {
  color: var(--color-accent);
  font-weight: 600;
}

.newsletter-box {
  background-color: var(--color-surface);
  border: 1px solid var(--color-border);
  border-radius: var(--radius-lg);
  padding: var(--space-6);
}

.newsletter-box__title {
  font-size: 1.125rem;
  font-weight: 600;
  color: var(--color-text);
  margin-bottom: var(--space-2);
}

.newsletter-box__text {
  font-size: 0.875rem;
  color: var(--color-text-body);
  line-height: 1.50;
  margin-bottom: var(--space-4);
}

.newsletter-box .form__input {
  margin-bottom: var(--space-3);
}

/* --- Species Card --- */
.species-card {
  display: grid;
  grid-template-columns: 1fr 1fr;
  gap: 0;
  background-color: var(--color-bg);
  border: 1px solid var(--color-border);
  border-radius: var(--radius-lg);
  overflow: hidden;
}

.species-card:nth-child(even) {
  direction: rtl;
}

.species-card:nth-child(even) > * {
  direction: ltr;
}

.species-card__image {
  background-color: var(--color-surface);
  display: flex;
  align-items: center;
  justify-content: center;
  min-height: 320px;
  font-size: 4rem;
  color: var(--color-text-muted);
}

.species-card__content {
  padding: var(--space-8);
  display: flex;
  flex-direction: column;
  justify-content: center;
}

.species-card__label {
  font-size: 0.75rem;
  font-weight: 600;
  text-transform: uppercase;
  letter-spacing: 0.5px;
  color: var(--color-accent);
  margin-bottom: var(--space-3);
}

.species-card__title {
  font-family: var(--font-display);
  font-size: 2rem;
  font-weight: 600;
  color: var(--color-text);
  margin-bottom: var(--space-4);
}

.species-card__text {
  font-size: 1rem;
  line-height: 1.50;
  color: var(--color-text-body);
  margin-bottom: var(--space-5);
}

.species-card__list {
  list-style: none;
  margin-bottom: var(--space-5);
}

.species-card__list li {
  font-size: 0.9375rem;
  color: var(--color-text-body);
  padding: var(--space-2) 0;
  border-bottom: 1px solid var(--color-surface);
  display: flex;
  align-items: center;
  gap: var(--space-2);
}

.species-card__list li::before {
  content: "✓";
  color: var(--color-accent);
  font-weight: 700;
  font-size: 0.75rem;
}

/* --- Service Section --- */
.service-section {
  padding: var(--space-12) 0;
  border-bottom: 1px solid var(--color-border);
}

.service-section:last-child {
  border-bottom: none;
}

.service-section__header {
  margin-bottom: var(--space-8);
}

.service-section__icon {
  display: inline-flex;
  align-items: center;
  justify-content: center;
  width: 56px;
  height: 56px;
  background-color: var(--color-surface);
  border: 1px solid var(--color-border);
  border-radius: var(--radius-lg);
  font-size: 1.5rem;
  margin-bottom: var(--space-4);
}

.service-section__title {
  font-family: var(--font-display);
  font-size: 2rem;
  font-weight: 600;
  color: var(--color-text);
  margin-bottom: var(--space-3);
}

.service-section__text {
  font-size: 1.0625rem;
  line-height: 1.50;
  color: var(--color-text-body);
  max-width: 720px;
}

/* --- Industry Section --- */
.industry-section {
  display: grid;
  grid-template-columns: 1fr 1fr;
  gap: var(--space-10);
  padding: var(--space-12) 0;
  border-bottom: 1px solid var(--color-border);
  align-items: center;
}

.industry-section:last-child {
  border-bottom: none;
}

.industry-section:nth-child(even) {
  direction: rtl;
}

.industry-section:nth-child(even) > * {
  direction: ltr;
}

.industry-section__visual {
  background-color: var(--color-surface);
  border: 1px solid var(--color-border);
  border-radius: var(--radius-lg);
  display: flex;
  align-items: center;
  justify-content: center;
  min-height: 280px;
  font-size: 3rem;
}

.industry-section__content {}

.industry-section__label {
  font-size: 0.75rem;
  font-weight: 600;
  text-transform: uppercase;
  letter-spacing: 0.5px;
  color: var(--color-accent);
  margin-bottom: var(--space-3);
}

.industry-section__title {
  font-size: 1.75rem;
  font-weight: 600;
  color: var(--color-text);
  margin-bottom: var(--space-4);
}

.industry-section__text {
  font-size: 1rem;
  line-height: 1.55;
  color: var(--color-text-body);
  margin-bottom: var(--space-5);
}

.industry-section__tests {
  list-style: none;
}

.industry-section__tests li {
  font-size: 0.9375rem;
  color: var(--color-text-body);
  padding: var(--space-2) 0;
  display: flex;
  align-items: center;
  gap: var(--space-2);
}

.industry-section__tests li::before {
  content: "→";
  color: var(--color-accent);
  font-weight: 700;
}

/* --- CTA Banner --- */
.cta-banner {
  background-color: var(--color-dark);
  color: var(--color-light);
  padding: var(--space-16) 0;
  text-align: center;
}

.cta-banner__title {
  font-family: var(--font-display);
  font-size: 2.5rem;
  font-weight: 500;
  line-height: 0.95;
  color: var(--color-light);
  margin-bottom: var(--space-4);
}

.cta-banner__text {
  font-size: 1.125rem;
  color: var(--color-border);
  max-width: 560px;
  margin: 0 auto var(--space-8);
  line-height: 1.50;
}

.cta-banner__actions {
  display: flex;
  gap: var(--space-4);
  justify-content: center;
  flex-wrap: wrap;
}

/* --- Compliance Cards --- */
.compliance-card {
  background-color: var(--color-bg);
  border: 1px solid var(--color-border);
  border-radius: var(--radius-lg);
  padding: var(--space-6);
}

.compliance-card__badge {
  display: inline-flex;
  align-items: center;
  justify-content: center;
  width: 48px;
  height: 48px;
  background-color: var(--color-surface);
  border: 1px solid var(--color-border);
  border-radius: var(--radius-md);
  font-size: 0.6875rem;
  font-weight: 700;
  color: var(--color-text);
  margin-bottom: var(--space-4);
}

.compliance-card__title {
  font-size: 1.25rem;
  font-weight: 600;
  color: var(--color-text);
  margin-bottom: var(--space-3);
}

.compliance-card__text {
  font-size: 0.9375rem;
  line-height: 1.55;
  color: var(--color-text-body);
}

/* --- Team --- */
.team-card {
  text-align: center;
}

.team-card__photo {
  width: 160px;
  height: 160px;
  border-radius: 50%;
  background-color: var(--color-surface);
  border: 1px solid var(--color-border);
  margin: 0 auto var(--space-4);
  display: flex;
  align-items: center;
  justify-content: center;
  font-size: 2.5rem;
  color: var(--color-text-muted);
  overflow: hidden;
}

.team-card__name {
  font-size: 1.125rem;
  font-weight: 600;
  color: var(--color-text);
  margin-bottom: 2px;
}

.team-card__role {
  font-size: 0.875rem;
  color: var(--color-accent);
  font-weight: 500;
  margin-bottom: var(--space-3);
}

.team-card__bio {
  font-size: 0.875rem;
  line-height: 1.55;
  color: var(--color-text-body);
  max-width: 280px;
  margin: 0 auto;
}

/* --- Testimonial --- */
.testimonial {
  background-color: var(--color-bg-alt);
  border: 1px solid var(--color-border);
  border-radius: var(--radius-lg);
  padding: var(--space-8);
  position: relative;
}

.testimonial__quote {
  font-family: var(--font-editorial);
  font-size: 1.5rem;
  font-weight: 300;
  line-height: 1.40;
  letter-spacing: -0.5px;
  color: var(--color-text);
  margin-bottom: var(--space-6);
  font-style: italic;
}

.testimonial__quote::before {
  content: "\201C";
  font-size: 3rem;
  color: var(--color-accent);
  line-height: 0;
  vertical-align: -0.4em;
  margin-right: 4px;
}

.testimonial__author {
  font-size: 0.9375rem;
  font-weight: 600;
  color: var(--color-text);
}

.testimonial__company {
  font-size: 0.8125rem;
  color: var(--color-text-muted);
}

/* --- Checklist --- */
.checklist {
  list-style: none;
}

.checklist li {
  padding: var(--space-3) 0;
  border-bottom: 1px solid var(--color-surface);
  font-size: 1rem;
  color: var(--color-text-body);
  display: flex;
  align-items: flex-start;
  gap: var(--space-3);
  line-height: 1.50;
}

.checklist__check {
  display: inline-flex;
  align-items: center;
  justify-content: center;
  flex-shrink: 0;
  width: 22px;
  height: 22px;
  background-color: var(--color-accent);
  color: var(--color-light);
  border-radius: 50%;
  font-size: 0.6875rem;
  font-weight: 700;
  margin-top: 1px;
}

/* --- Map Placeholder --- */
.map-placeholder {
  width: 100%;
  height: 320px;
  background-color: var(--color-surface);
  border: 1px solid var(--color-border);
  border-radius: var(--radius-lg);
  display: flex;
  align-items: center;
  justify-content: center;
  color: var(--color-text-muted);
  font-size: 1rem;
}

/* --- Case Study Detail --- */
.case-study-detail__meta {
  display: flex;
  gap: var(--space-6);
  margin-bottom: var(--space-8);
  flex-wrap: wrap;
}

.case-study-detail__meta-item {
  font-size: 0.875rem;
}

.case-study-detail__meta-label {
  font-weight: 600;
  color: var(--color-text);
  display: block;
  margin-bottom: 2px;
}

.case-study-detail__meta-value {
  color: var(--color-text-body);
}

.case-study-detail__section {
  margin-bottom: var(--space-10);
}

.case-study-detail__section-title {
  font-size: 1.5rem;
  font-weight: 600;
  color: var(--color-text);
  margin-bottom: var(--space-4);
  padding-bottom: var(--space-3);
  border-bottom: 1px solid var(--color-border);
}

.case-study-detail__section-text {
  font-size: 1.0625rem;
  line-height: 1.65;
  color: var(--color-text-body);
}

/* --- Contact Info --- */
.contact-info {
  list-style: none;
}

.contact-info__item {
  display: flex;
  gap: var(--space-4);
  padding: var(--space-4) 0;
  border-bottom: 1px solid var(--color-surface);
  align-items: flex-start;
}

.contact-info__icon {
  display: flex;
  align-items: center;
  justify-content: center;
  flex-shrink: 0;
  width: 40px;
  height: 40px;
  background-color: var(--color-surface);
  border-radius: var(--radius-md);
  font-size: 1.125rem;
}

.contact-info__label {
  font-size: 0.8125rem;
  font-weight: 600;
  text-transform: uppercase;
  letter-spacing: 0.3px;
  color: var(--color-text-muted);
  margin-bottom: 2px;
}

.contact-info__value {
  font-size: 1rem;
  color: var(--color-text);
}

.contact-info__value a {
  color: var(--color-text);
}

/* --- Footer --- */
.footer {
  background-color: var(--color-dark);
  color: var(--color-light);
  padding: var(--space-12) 0 var(--space-8);
}

.footer__logo {
  display: flex;
  align-items: center;
  gap: var(--space-2);
  margin-bottom: 0;
}

.footer__logo-text {
  font-family: var(--font-display);
  font-size: 1.25rem;
  font-weight: 700;
  color: var(--color-light);
}

.footer__badges {
  margin-top: var(--space-5);
  display: flex;
  gap: var(--space-3);
}

.footer__inner {
  display: grid;
  grid-template-columns: 2fr 1fr 1fr 1fr;
  gap: var(--space-10);
  max-width: var(--container-max);
  margin: 0 auto;
  padding: 0 var(--container-padding);
}

.footer__brand-text {
  font-size: 0.9375rem;
  color: var(--color-border);
  line-height: 1.55;
  margin-top: var(--space-4);
  max-width: 320px;
}

.footer__heading {
  font-size: 0.8125rem;
  font-weight: 600;
  text-transform: uppercase;
  letter-spacing: 0.5px;
  color: var(--color-light);
  margin-bottom: var(--space-4);
}

.footer__links {
  list-style: none;
}

.footer__link {
  display: block;
  padding: var(--space-1) 0;
  font-size: 0.9375rem;
  color: var(--color-border);
  text-decoration: none;
  transition: color 0.15s ease;
}

.footer__link:hover {
  color: var(--color-light);
  text-decoration: none;
}

.footer__bottom {
  max-width: var(--container-max);
  margin: var(--space-12) auto 0;
  padding: var(--space-6) var(--container-padding) 0;
  border-top: 1px solid rgba(197, 192, 177, 0.2);
  display: flex;
  justify-content: space-between;
  align-items: center;
  font-size: 0.8125rem;
  color: var(--color-text-muted);
  flex-wrap: wrap;
  gap: var(--space-4);
}

.footer__bottom-links {
  display: flex;
  gap: var(--space-6);
}

.footer__bottom-links a {
  color: var(--color-text-muted);
  text-decoration: none;
}

.footer__bottom-links a:hover {
  color: var(--color-light);
}

/* --- Image Fills --- */
.species-card__image img,
.industry-section__visual img {
  width: 100%;
  height: 100%;
  object-fit: cover;
}

.team-card__photo img {
  width: 100%;
  height: 100%;
  object-fit: cover;
}

/* --- Accreditation Icon --- */
.accred-icon {
  display: flex;
  align-items: center;
  justify-content: center;
  width: 64px;
  height: 64px;
  margin: 0 auto var(--space-4);
  background-color: var(--color-bg);
  border: 2px solid var(--color-accent);
  border-radius: 50%;
  color: var(--color-accent);
  font-size: 0.75rem;
  font-weight: 800;
  letter-spacing: 0.5px;
  position: relative;
}

.accred-icon::after {
  content: "";
  position: absolute;
  inset: 3px;
  border: 1px solid var(--color-accent);
  border-radius: 50%;
  opacity: 0.4;
}

.accred-card {
  background-color: var(--color-bg);
  border: 1px solid var(--color-border);
  border-radius: var(--radius-lg);
  padding: var(--space-6) var(--space-5);
  text-align: center;
  transition: border-color 0.2s ease;
}

.accred-card:hover {
  border-color: var(--color-accent);
}

.accred-card__name {
  font-size: 0.9375rem;
  font-weight: 600;
  color: var(--color-text);
  margin-bottom: 4px;
}

.accred-card__desc {
  font-size: 0.8125rem;
  color: var(--color-text-muted);
  line-height: 1.4;
}

/* --- Quote Card (dark variant) --- */
.quote-card {
  background-color: var(--color-dark);
  color: var(--color-light);
  border-color: var(--color-dark);
}

.quote-card .card__title {
  color: var(--color-light);
}

.quote-card .card__text {
  color: var(--color-border);
}

/* --- Utility --- */
.btn--block {
  width: 100%;
}

/* --- Scroll Animations --- */
.fade-in {
  opacity: 0;
  transform: translateY(24px);
  transition: opacity 0.6s ease, transform 0.6s ease;
}

.fade-in--visible {
  opacity: 1;
  transform: translateY(0);
}

.fade-in--delay-1 { transition-delay: 0.1s; }
.fade-in--delay-2 { transition-delay: 0.2s; }
.fade-in--delay-3 { transition-delay: 0.3s; }
.fade-in--delay-4 { transition-delay: 0.4s; }

/* --- Utility --- */
.visually-hidden {
  position: absolute;
  width: 1px;
  height: 1px;
  padding: 0;
  margin: -1px;
  overflow: hidden;
  clip: rect(0, 0, 0, 0);
  white-space: nowrap;
  border: 0;
}

.divider {
  border: none;
  border-top: 1px solid var(--color-border);
  margin: var(--space-8) 0;
}

/* --- Responsive --- */

/* Tablet Large and below */
@media (max-width: 991px) {
  :root {
    --container-padding: 20px;
  }

  .nav__links {
    display: none;
  }

  .nav__cta.desktop-only {
    display: none;
  }

  .nav__mobile-toggle {
    display: flex;
  }

  .hero--home .hero__title {
    font-size: 3.5rem;
  }

  .hero__title {
    font-size: 2.5rem;
  }

  .section {
    padding: var(--space-12) 0;
  }

  .section--lg {
    padding: var(--space-14) 0;
  }

  .grid--4 {
    grid-template-columns: repeat(2, 1fr);
  }

  .footer__inner {
    grid-template-columns: 1fr 1fr;
    gap: var(--space-8);
  }

  .blog-layout {
    grid-template-columns: 1fr;
  }

  .blog-sidebar {
    position: static;
  }

  .industry-section {
    grid-template-columns: 1fr;
  }

  .industry-section:nth-child(even) {
    direction: ltr;
  }

  .industry-section__visual {
    min-height: 200px;
  }

  .species-card {
    grid-template-columns: 1fr;
  }

  .species-card:nth-child(even) {
    direction: ltr;
  }

  .species-card__image {
    min-height: 220px;
  }

  .card--horizontal {
    grid-template-columns: 1fr;
  }

  .card--horizontal .card__image {
    border-bottom: 1px solid var(--color-border);
    border-right: none;
    aspect-ratio: 16 / 10;
  }

  .stats {
    gap: var(--space-8);
  }

  .case-study-detail__meta {
    gap: var(--space-4);
  }
}

/* Tablet and below */
@media (max-width: 768px) {
  .hero--home .hero__title {
    font-size: 2.75rem;
  }

  .hero__title {
    font-size: 2.25rem;
  }

  .section-heading {
    font-size: 2.25rem;
  }

  .grid--3 {
    grid-template-columns: repeat(2, 1fr);
  }

  .grid--2 {
    grid-template-columns: 1fr;
  }

  .stats {
    justify-content: center;
    gap: var(--space-6);
  }

  .stats__number {
    font-size: 2.5rem;
  }

  .trust-bar {
    gap: var(--space-6);
  }

  .form__row {
    grid-template-columns: 1fr;
  }

  .cta-banner__title {
    font-size: 2rem;
  }

  .service-section__title {
    font-size: 1.5rem;
  }
}

/* Mobile */
@media (max-width: 600px) {
  :root {
    --container-padding: 16px;
  }

  .hero {
    padding: var(--space-10) 0 var(--space-8);
  }

  .hero--home {
    padding: var(--space-12) 0 var(--space-10);
  }

  .hero--home .hero__title {
    font-size: 2.25rem;
  }

  .hero__title {
    font-size: 2rem;
  }

  .hero__subtitle {
    font-size: 1.0625rem;
  }

  .hero__actions {
    flex-direction: column;
  }

  .hero__actions .btn {
    width: 100%;
    text-align: center;
  }

  .section {
    padding: var(--space-10) 0;
  }

  .section-heading {
    font-size: 1.75rem;
  }

  .grid--3,
  .grid--4 {
    grid-template-columns: 1fr;
  }

  .stats {
    flex-direction: column;
    align-items: center;
    gap: var(--space-6);
  }

  .trust-bar {
    flex-direction: column;
    gap: var(--space-4);
  }

  .footer__inner {
    grid-template-columns: 1fr;
    gap: var(--space-8);
  }

  .footer__bottom {
    flex-direction: column;
    text-align: center;
  }

  .cta-banner {
    padding: var(--space-12) 0;
  }

  .cta-banner__title {
    font-size: 1.75rem;
  }

  .cta-banner__actions {
    flex-direction: column;
    align-items: center;
  }

  .filter-bar {
    gap: var(--space-1);
  }

  .filter-btn {
    font-size: 0.8125rem;
    height: 32px;
    padding: 0 12px;
  }

  .card__title {
    font-size: 1.25rem;
  }

  .card__body {
    padding: var(--space-5);
  }

  .species-card__content {
    padding: var(--space-6);
  }

  .editorial-heading {
    font-size: 2rem;
  }

  .editorial-sub {
    font-size: 1.75rem;
  }

  .sub-heading-lg {
    font-size: 1.75rem;
  }

  .sub-heading {
    font-size: 1.5rem;
  }

  .sub-heading-md {
    font-size: 1.25rem;
  }

  .display-hero-xl {
    font-size: 2.5rem;
  }

  .display-hero {
    font-size: 2rem;
  }

  .display-hero-sm {
    font-size: 1.75rem;
  }
}

/* Small Mobile */
@media (max-width: 450px) {
  .hero--home .hero__title {
    font-size: 1.875rem;
  }

  .hero__title {
    font-size: 1.75rem;
  }

  .section-heading {
    font-size: 1.5rem;
  }

  .stats__number {
    font-size: 2rem;
  }

  .breadcrumb {
    font-size: 0.8125rem;
  }
}